Simulate and analyze genetic drift in populations. Understand how random chance affects allele frequencies over generations.
Genetic drift is the change in the frequency of an existing gene variant (allele) in a population due to random sampling of organisms. The alleles in the offspring generation are a random sample of those in the parent generation, and chance plays a role in determining whether a given individual survives and reproduces.
Key Insight: Genetic drift has a larger effect in small populations and can lead to the loss of genetic variation, fixation of alleles, and divergence between populations.
Wright-Fisher Model: The classic model of genetic drift with discrete generations. Each generation is formed by random sampling of alleles from the previous generation. This model assumes constant population size and random mating.
Moran Model: A model with overlapping generations where individuals reproduce and die one at a time. This model more closely resembles populations with continuous reproduction rather than discrete generations.
The effective population size (Nₑ) is the size of an idealized population that would experience the same amount of genetic drift as the actual population. Real populations often have Nₑ smaller than the census population size due to:
Conservation Importance: Effective population size is crucial for conservation genetics. Small Nₑ increases the rate of genetic drift, leading to loss of genetic diversity and increased inbreeding, which can reduce population viability.
| Property | Formula | Explanation |
|---|---|---|
| Fixation Probability | Pfix = p | For a neutral allele, probability of fixation equals its current frequency |
| Rate of Heterozygosity Loss | Ht = H0(1 - 1/(2Ne))t | Heterozygosity decreases over time due to genetic drift |
| Variance in Allele Frequency | σ2 = p(1-p)/(2Ne) | Variance in allele frequency change per generation |
| Time to Fixation | T ≈ 4Ne generations | Average time for a neutral allele to become fixed |
Hunted to near extinction in the 19th century, the population rebounded from about 20 individuals, resulting in extremely low genetic diversity.
This isolated population founded by 15 people has higher rates of certain genetic diseases due to founder effect.
Several genetic disorders are more common in Amish populations due to founder effect and genetic drift.